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Environmental complexity constrains evolutionary adaptation across taxa

Zilio, G.; Aubin, E.; Bedhomme, S.; Bolick, L.; Bravo, I. G.; Bruand, C.; Capela, D.; Challe, M.; Charriere, G. M.; Courtay, G.; Devillez, M.-A.; Elmaleh, F.; Fereol, S.; Froissart, R.; Givens, J.; Gougat-Barbera, C.; Govaert, L.; Guidot, A.; Hamet, J.; Huet, M.; Hummer, P.; Jacob, S.; Kaltz, O.; Krasovec, M.; Legrand, D.; Martin, G.; Nidelet, T.; Orcel, D.; Philippe, H.; Piganeau, G.; Przybylska, M. S.; Remigi, P.; Sauviac, L.; Schneider-Nettstrater, F.; Segond, D.; Serre, C.; Sicard, D.; Silveira, J. G. C.; Tonnabel, J.; Vasseur, F.; Vedrenne, A.; Vidal, E.; Violle, C.; Wenzel, M. K.; Fronho

2026-04-29 evolutionary biology
10.64898/2025.12.22.695988 bioRxiv
Show abstract

Complex environments combining multiple stressors are the new norm worldwide. Adaptive evolution will be critical to population persistence under these combined challenges, but how environmental complexity affects the pace of evolution remains poorly understood. Using a meta-experimental evolution approach, we exposed 14 species, from bacteria to unicellular eukaryotes and plants, to single stressors and their pairwise combinations for multiple generations, while keeping the overall stress level comparable. Populations evolving under combined stressors had lower fitness increase in the selective environments, higher fitness reductions in the control environment, and shallower relation between initial maladaptation and fitness gain, than under single stressors. However, these responses varied with species and stressor type. Accounting for such constraints on evolutionary dynamics should prove crucial for the management of biodiversity.

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